Thursday, May 28, 2009

Over Population

By: Aldous Huxley [1958]

In the light of what we have recently learned about animal behavior in general, and human behavior in particular, it has become clear that control through the punishment of undesirable behavior is less effec­tive, in the long run, than control through the rein­forcement of desirable behavior by rewards, and that government through terror works on the whole less well than government through the non-violent manip­ulation of the environment and of the thoughts and feelings of individual men, women and children. Pun­ishment temporarily puts a stop to undesirable behav­ior, but does not permanently reduce the victim's tend­ency to indulge in it. Moreover, the psycho-physical by-products of punishment may be just as undesirable as the behavior for which an individual has been pun­ished. Psychotherapy is largely concerned with the de­bilitating or anti-social consequences of past punish­ments.

On the first Christmas Day the population of our planet was about two hundred and fifty millions -- less than half the population of modern China. Sixteen cen­turies later, when the Pilgrim Fathers landed at Plym­outh Rock, human numbers had climbed to a little more than five hundred millions. By the time of the signing of the Declaration of Independence, world pop­ulation had passed the seven hundred million mark. In 1931, when I was writing Brave New World, it stood at just under two billions. Today, only twenty-seven years later, there are two billion eight hundred million of us. And tomorrow -- what? Penicillin, DDT and clean water are cheap commodities, whose effects on public health are out of all proportion to their cost. Even the poorest government is rich enough to provide its subjects with a substantial measure of death con­trol. Birth control is a very different matter. Death control is something which can be provided for a whole people by a few technicians working in the pay of a benevolent government. Birth control depends on the co-operation of an entire people. It must be practiced by countless individuals, from whom it demands more intelligence and will power than most of the world's teeming illiterates possess, and (where chemical or me­chanical methods of contraception are used) an ex­penditure of more money than most of these millions can now afford. Moreover, there are nowhere any reli­gious traditions in favor of unrestricted death, whereas religious and social traditions in favor of un­restricted reproduction are widespread. For all these reasons, death control is achieved very easily, birth control is achieved with great difficulty. Death rates have therefore fallen in recent years with startling suddenness. But birth rates have either remained at their old high level or, if they have fallen, have fallen very little and at a very slow rate. In consequence, human numbers are now increasing more rapidly than at any time in the history of the species.

Moreover, the yearly increases are themselves in­creasing. They increase regularly, according to the rules of compound interest; and they also increase irregularly with every application, by a technologically backward society of the principles of Public Health. At the present time the annual increase in world pop­ulation runs to about forty-three millions. This means that every four years mankind adds to its numbers the equivalent of the present population of the United States, every eight and a half years the equivalent of the present population of India. At the rate of increase prevailing between the birth of Christ and the death of Queen Elizabeth I, it took sixteen centuries for the population of the earth to double. At the present rate it will double in less than half a century. And this fantastically rapid doubling of our numbers will be taking place on a planet whose most desirable and pro­ductive areas are already densely populated, whose soils are being eroded by the frantic efforts of bad farmers to raise more food, and whose easily available mineral capital is being squandered with the reckless extravagance of a drunken sailor getting rid of his accumulated pay.

As large and increasing numbers press more heavily upon available resources, the economic position of the society undergoing this ordeal becomes ever more precarious. This is especially true of those underdeveloped regions, where a sudden lowering of the death rate by means of DDT, penicillin and clean water has not been accompanied by a corresponding fall in the birth rate. In parts of Asia and in most of Central and South America populations are increasing so fast that they will double themselves in little more than twenty years. If the production of food and manufactured arti­cles, of houses, schools and teachers, could be in­creased at a greater rate than human numbers, it would be possible to improve the wretched lot of those who live in these underdeveloped and over-populated countries. But unfortunately these countries lack not merely agricultural machinery and an industrial plant capable of turning out this machinery, but also the capital required to create such a plant. Capital is what is left over after the primary needs of a population have been satisfied. But the primary needs of most of the people in underdeveloped countries are never fully satisfied. At the end of each year almost nothing is left over, and there is therefore almost no capital avail­able for creating the industrial and agricultural plant, by means of which the people's needs might be satisfied. Moreover, there is, in all these underdevel­oped countries, a serious shortage of the trained man­power without which a modern industrial and agricul­tural plant cannot be operated. The present educa­tional facilities are inadequate; so are the resources, financial and cultural, for improving the existing facili­ties as fast as the situation demands. Meanwhile the population of some of these underdeveloped countries is increasing at the rate of 3 per cent per annum.

Whenever the economic life of a nation becomes pre­carious, the central government is forced to assume additional responsibilities for the general welfare. It must work out elaborate plans for dealing with a criti­cal situation; it must impose ever greater restrictions upon the activities of its subjects; and if, as is very likely, worsening economic conditions result in polit­ical unrest, or open rebellion, the central government must intervene to preserve public order and its own authority. More and more power is thus concentrated in the hands of the executives and their bureaucratic managers. But the nature of power is such that even those who have not sought it, but have had it forced upon them, tend to acquire a taste for more. "Lead us not into temptation," we pray -- and with good reason; for when human beings are tempted too enticingly or too long, they generally yield. A democratic constitu­tion is a device for preventing the local rulers from yielding to those particularly dangerous temptations that arise when too much power is concentrated in too few hands. Such a constitution works pretty well where, as in Britain or the United States, there is a traditional respect for constitutional procedures. Where the republican or limited monarchical tradition is weak, the best of constitutions will not prevent ambi­tious politicians from succumbing with glee and gusto to the temptations of power. And in any country where numbers have begun to press heavily upon avail­able resources, these temptations cannot fail to arise. Over-population leads to economic insecurity and so­cial unrest. Unrest and insecurity lead to more con­trol by central governments and an increase of their power. In the absence of a constitutional tradition, this increased power will probably be exercised in a dictatorial fashion. Even if Communism had never been invented, this would be likely to happen. But Com­munism has been invented. Given this fact, the proba­bility of over-population leading through unrest to dic­tatorship becomes a virtual certainty. It is a pretty safe bet that, twenty years from now, all the world's over-populated and underdeveloped countries will be under some form of totalitarian rule -- probably by the Communist party.

How will this development affect the over-populated, but highly industrialized and still democratic coun­tries of Europe? If the newly formed dictatorships were hostile to them, and if the normal flow of raw materials from the underdeveloped countries were de­liberately interrupted, the nations of the West would find themselves in a very bad way indeed. Their in­dustrial system would break down, and the highly de­veloped technology, which up till now has permitted them to sustain a population much greater than that which could be supported by locally available resources, would no longer protect them against the consequences of having too many people in too small a territory. If this should happen, the enormous powers forced by unfavorable conditions upon central govern­ments may come to be used in the spirit of totalitarian dictatorship.

The United States is not at present an over-popu­lated country. If, however, the population continues to increase at the present rate (which is higher than that of India's increase, though happily a good deal lower than the rate now current in Mexico or Guatemala), the problem of numbers in relation to available resources might well become troublesome by the begin­ning of the twenty-first century. For the moment over­population is not a direct threat to the personal free­dom of Americans. It remains, however, an indirect threat, a menace at one remove. If over-population should drive the underdeveloped countries into totali­tarianism, and if these new dictatorships should ally themselves with Russia, then the military position of the United States would become less secure and the preparations for defense and retaliation would have to be intensified. But liberty, as we all know, cannot flour­ish in a country that is permanently on a war footing, or even a near-war footing. Permanent crisis justifies permanent control of everybody and everything by the agencies of the central government. And permanent crisis is what we have to expect in a world in which over-population is producing a state of things, in which dictatorship under Communist auspices becomes almost inevitable.

Tuesday, May 26, 2009

Human Evolution

The Evolution of the Human

The universe is constructed from a multitude of various materials. It is dynamic in form and shape due to a multitude of various processes and interactions between these materials. To the human, however, in his need to establish his place and purpose in the universe, the most important material is biological and the most important process is evolution, far it is only here that the human can learn to understand himself, an understanding that is vital to his survival.

Wise men, psychologists, philosophers and theologians have surmised and conjectured about the human over the centuries, and still do, but the truth about the human may be found only through factual knowledge. That factual knowledge lies in a process called evolution. The human is what evolution made him.

History of Man

SPECIES TIME PERIOD
Ardipithicus ramidus 5 to 4 million years ago
Australopithecus anamensis 4.2 to 3.9 million years ago
Australopithecus afarensis 4 to 2.7 million years ago
Australopithecus africanus 3 to 2 million years ago
Australopithecus robustus 2.2 to 1.6 million years ago
Homo habilis 2.2 to 1.6 million years ago
Homo erectus 2.0 to 0.4 million years ago
Homo sapiens archaic 400 to 200 thousand years ago
Homo sapiens neandertalensis 200 to 30 thousand years ago
Homo sapiens sapiens 200 thousand years ago to present

The times of existence of the various hominid shown in the chart above are based on dated fossil remains. Each species may have existed earlier and/or later than shown, but fossil proof has not been discovered yet. There is also dispute concerning many overlapping species, for example, the overlap between Homo habilis and Homo erectus. It could well be that the two are continuing examples of the same species. The same dispute exists with Homo erectus, Homo sapiens archaic and homo sapiens sapiens. If all species have been discovered and the lineage of man lies within them, the most probable lineage would include all but the robust Australopithecines and the neandertal.

The following chronology is abbreviated:

The earliest fossil hominid, Ardipithecus ramidus, is a recent discovery. It is dated at 4.4 million years ago. The remains are incomplete but enough is available to suggest it was bipedal and about 4 feet tall. Other fossils were found with the ramidus fossil which would suggest that ramidus was a forest dweller. A new skeleton was recently discovered which is about 45% complete. It is now being studied.

A new species, Australopithecus anamensis, was named in 1995. It was found in Allia Bay in Kenya. Anamensis lived between 4.2 and 3.9 million years ago. Its body showed advanced bipedal features, but the skull closely resembled the ancient apes.

Australopithecus afarensis lived between 3.9 and 3.0 million years ago. It retained the apelike face with a sloping forehead, a distinct ridge over the eyes, flat nose and a chinless lower jaw. It had a brain capacity of about 450 cc. It was between 3'6" and 5' tall. It was fully bipedal and the thickness of its bones showed that it was quite strong. Its build (ratio of weight to height) was about the same as the modern human but its head and face were proportionately much larger. This larger head with powerful jaws is a feature of all species prior to Homo sapiens sapiens.

Australopithecus africanus was quite similar to afarensis and lived between three and two million years ago. It was also bipedal, but was slightly larger in body size. Its brain size was also slightly larger, ranging up to 500 cc. The brain was not advanced enough for speech. The molars were a little larger than in afarensis and much larger than modern human. This hominid was a herbivore and ate tough, hard to chew, plants. The shape of the jaw was now like the human.

Australopithecus aethiopicus lived between 2.6 and 2.3 million years ago. This species is probably an ancestor of the robustus and boisei. This hominid ate a rough and hard to chew diet. He had huge molars and jaws and a large sagittal crest. A sagittal crest is a bony ridge on the skull extending from the forehead to the back of the head. Massive chewing muscles were anchored to this crest. See the opening picture of an early Homo habilis for an example. Brain sizes were still about 500cc, with no indication of speech functions.

Australopithecus robustus lived between two and 1.5 million years ago. It had a body similar to that of africanus, but a larger and more massive skull and teeth. Its huge face was flat and with no forehead. It had large brow ridges and a sagittal crest. Brain size was up to 525cc with no indication of speech capability.

Australopithecus boisei lived between 2.1 and 1.1 million years ago. It was quite similar to robustus, but with an even more massive face. It had huge molars, the larger measuring 0.9 inches across. The brain size was about the same as robustus. Some authorities believe that robustus and boisei are variants of the same species.

Homo habilis was called the handy man because tools were found with his fossil remains. This species existed between 2.4 and 1.5 million years ago. The brain size in earlier fossil specimens was about 500cc but rose to 800cc toward the end of the species life period. The species brain shape shows evidence that some speech had developed. Habilis was about 5' tall and weighed about 100 pounds. Some scientists believe that habilis is not a separate species and should be carried either as a later Australopithecine or an early Homo erectus. It is possible that early examples are in one species group and later examples in the other.

Homo erectus lived between 1.8 million and 300,000 years ago. It was a successful species for a million and a half years. Early examples had a 900cc brain size on the average. The brain grew steadily during its reign. Toward the end its brain was almost the same size as modern man, at about 1200cc. The species definitely had speech. Erectus developed tools, weapons and fire and learned to cook his food. He traveled out of Africa into China and Southeast Asia and developed clothing for northern climates. He turned to hunting for his food. Only his head and face differed from modern man. Like habilis, the face had massive jaws with huge molars, no chin, thick brow ridges, and a long low skull. Though proportioned the same, he was sturdier in build and much stronger than the modern human.

Homo sapiens (archaic) provides the bridge between erectus and Homo sapiens sapiens during the period 200,000 to 500,000 years ago. Many skulls have been found with features intermediate between the two. Brain averaged about 1200cc and speech was indicated. Skulls are more rounded and with smaller features. Molars and brow ridges are smaller. The skeleton shows a stronger build than modern human but was well proportioned.

Homo sapiens neandertalensis lived in Europe and the Mideast between 150,000 and 35,000 years ago. Neandertals coexisted with H.sapiens (archaic) and early H.sapiens sapiens. It is not known whether he was of the same species and disappeared into the H.sapiens sapiens gene pool or he may have been crowded out of existence (killed off) by the H.sapien sapien. Recent DNA studies have indicated that the neandertal was an entirely different species and did not merge into the H. sapiens sapiens gene pool. Brain sizes averaged larger than modern man at about 1450cc but the head was shaped differently, being longer and lower than modern man. His nose was large and was different from modern man in structure. He was a massive man at about 5'6" tall with an extremely heavy skeleton that showed attachments for massive muscles. He was far stronger than modern man. His jaw was massive and he had a receding forehead, like erectus.

Homo sapiens sapiens first appeared about 120,000 years ago. Modern humans have an average brain size of about 1350 cc.

excerpt take from http://www.onelife.com/evolve/manev.html

Fusion vs Fission

Fission

Nuclear fission is a nuclear reaction in which the nucleus of an atom splits into smaller parts, often producing free neutrons and lighter nuclei, which may eventually produce photons (in the form of gamma rays). Fission of heavy elements is an exothermic reaction which can release large amounts of energy both as electromagnetic radiation and as kinetic energy of the fragments (heating the bulk material where fission takes place). Fission is a form of nuclear transmutation because the resulting fragments are not the same element as the original atom.

Nuclear fission produces energy for nuclear power and to drive the explosion of nuclear weapons. Both uses are made possible because certain substances called nuclear fuels undergo fission when struck by free neutrons and in turn generate neutrons when they break apart. This makes possible a self-sustaining chain reaction that releases energy at a controlled rate in a nuclear reactor or at a very rapid uncontrolled rate in a nuclear weapon.

The amount of free energy contained in nuclear fuel is millions of times the amount of free energy contained in a similar mass of chemical fuel such as gasoline, making nuclear fission a very tempting source of energy; however, the products of nuclear fission are radioactive and remain so for significant amounts of time, giving rise to a nuclear waste problem. Concerns over nuclear waste accumulation and over the destructive potential of nuclear weapons may counterbalance the desirable qualities of fission as an energy source, and give rise to ongoing political debate over nuclear power.

Fusion

Nuclear fusion is the process by which multiple like-charged atomic nuclei join together to form a heavier nucleus. It is accompanied by the release or absorption of energy, which allows matter to enter a plasma state.

The fusion of two nuclei with lower mass than iron (which, along with nickel, has the largest binding energy per nucleon) generally releases energy while the fusion of nuclei heavier than iron absorbs energy; vice-versa for the reverse process, nuclear fission. In the simplest case of hydrogen fusion, two protons have to be brought close enough for their mutual electric repulsion to be overcome by the nuclear force and the subsequent release of energy.

Nuclear fusion occurs naturally in stars. Artificial fusion in human enterprises has also been achieved, although has not yet been completely controlled. Building upon the nuclear transmutation experiments of Ernest Rutherford done a few years earlier, fusion of light nuclei (hydrogen isotopes) was first observed by Mark Oliphant in 1932; the steps of the main cycle of nuclear fusion in stars were subsequently worked out by Hans Bethe throughout the remainder of that decade. Research into fusion for military purposes began in the early 1940s as part of the Manhattan Project, but was not successful until 1952. Research into controlled fusion for civilian purposes began in the 1950s, and continues to this day.

excerpts taken from wikipidea

Monday, May 25, 2009

Vacuum Fluctuations

Some interesting views on the vacuum fluctuations which began the universe
  • There are something like ten million million million million million million million million million million million million million million (1 with eighty [five] zeroes after it) particles in the region of the universe that we can observe. Where did they all come from? The answer is that, in quantum theory, particles can be created out of energy in the form of particle/antiparticle pairs. But that just raises the question of where the energy came from. The answer is that the total energy of the universe is exactly zero. The matter in the universe is made out of positive energy. However, the matter is all attracting itself by gravity. Two pieces of matter that are close to each other have less energy than the same two pieces a long way apart, because you have to expend energy to separate them against the gravitational force that is pulling them together. Thus, in a sense, the gravitational field has negative energy. In the case of a universe that is approximately uniform in space, one can show that this negative gravitational energy exactly cancels the positive energy represented by the matter. So the total energy of the universe is zero. (Hawking, 1988, 129) [thanks to Ross King for this quote]


  • There is a still more remarkable possibility, which is the creation of matter from a state of zero energy. This possibility arises because energy can be both positive and negative. The energy of motion or the energy of mass is always positive, but the energy of attraction, such as that due to certain types of gravitational or electromagnetic field, is negative. Circumstances can arise in which the positive energy that goes to make up the mass of newly-created particles of matter is exactly offset by the negative energy of gravity of electromagnetism. For example, in the vicinity of an atomic nucleus the electric field is intense. If a nucleus containing 200 protons could be made (possible but difficult), then the system becomes unstable against the spontaneous production of electron-positron pairs, without any energy input at all. The reason is that the negative electric energy can exactly offset the energy of their masses.

    In the gravitational case the situation is still more bizarre, for the gravitational field is only a spacewarp - curved space. The energy locked up in a spacewarp can be converted into particles of matter and antimatter. This occurs, for example, near a black hole, and was probably also the most important source of particles in the big bang. Thus, matter appears spontaneously out of empty space. The question then arises, did the primeval bang possess energy, or is the entire universe a state of zero energy, with the energy of all the material offset by negative energy of gravitational attraction?

    It is possible to settle the issue by a simple calculation. Astronomers can measure the masses of galaxies, their average separation, and their speeds of recession. Putting these numbers into a formula yields a quantity which some physicists have interpreted as the total energy of the universe. The answer does indeed come out to be zero within the observational accuracy. The reason for this distinctive result has long been a source of puzzlement to cosmologists. Some have suggested that there is a deep cosmic principle at work which requires the universe to have exactly zero energy. If that is so the cosmos can follow the path of least resistance, coming into existence without requiring any input of matter or energy at all. (Davies, 1983, 31-32)


  • Once our minds accept the mutability of matter and the new idea of the vacuum, we can speculate on the origin of the biggest thing we know - the universe. Maybe the universe itself sprang into existence out of nothingness - a gigantic vacuum fluctuation which we know today as the big bang. Remarkably, the laws of modern physics allow for this possibility. (Pagels, 1982, 247)


  • In general relativity, spacetime can be empty of matter or radiation and still contain energy stored in its curvature. Uncaused, random quantum fluctuations in a flat, empty, featureless spacetime can produce local regions with positive or negative curvature. This is called the "spacetime foam" and the regions are called "bubbles of false vacuum." Wherever the curvature is positive a bubble of false vacuum will, according to Einstein's equations, exponentially inflate. In 10-42 seconds the bubble will expand to the size of a proton and the energy within will be sufficient to produce all the mass of the universe.

    The bubbles start out with no matter, radiation, or force fields and maximum entropy. They contain energy in their curvature, and so are a "false vacuum." As they expand, the energy within increases exponentially. This does not violate energy conservation since the false vacuum has a negative pressure (believe me, this is all follows from the equations that Einstein wrote down in 1916) so the expanding bubble does work on itself.

    As the bubble universe expands, a kind of friction occurs in which energy is converted into particles. The temperature then drops and a series of spontaneous symmetry breaking processes occurs, as in a magnet cooled below the Curie point and a essentially random structure of the particles and forces appears. Inflation stops and we move into the more familiar big bang.

    The forces and particles that appear are more-or-less random, governed only by symmetry principles (like the conservation principles of energy and momentum) that are also not the product of design but exactly what one has in the absence of design.

    The so-called "anthropic coincidences," in which the particles and forces of physics seem to be "fine-tuned" for the production of Carbon-based life are explained by the fact that the spacetime foam has an infinite number of universes popping off, each different. We just happen to be in the one where the forces and particles lent themselves to the generation of carbon and other atoms with the complexity necessary to evolve living and thinking organisms. (Stenger, 1996)


  • Where did all the matter and radiation in the universe come from in the first place? Recent intriguing theoretical research by physicists such as Steven Weinberg of Harvard and Ya. B. Zel'dovich in Moscow suggest that the universe began as a perfect vacuum and that all the particles of the material world were created from the expansion of space...

    Think about the universe immediately after the Big Bang. Space is violently expanding with explosive vigor. Yet, as we have seen, all space is seething with virtual pairs of particles and antiparticles. Normally, a particle and anti-particle have no trouble getting back together in a time interval...short enough so that the conservation of mass is satisfied under the uncertainty principle. During the Big Bang, however, space was expanding so fast that particles were rapidly pulled away from their corresponding antiparticles. Deprived of the opportunity to recombine, these virtual particles had to become real particles in the real world. Where did the energy come from to achieve this materialization?

    Recall that the Big Bang was like the center of a black hole. A vast supply of gravitational energy was therefore associated with the intense gravity of this cosmic singularity. This resource provided ample energy to completely fill the universe with all conceivable kinds of particles and antiparticles. Thus, immediately after the Planck time, the universe was flooded with particles and antiparticles created by the violent expansion of space. (Kaufmann, 1985, 529-532)


  • ...the idea of a First Cause sounds somewhat fishy in light of the modern theory of quantum mechanics. According to the most commonly accepted interpretation of quantum mechanics, individual subatomic particles can behave in unpredictable ways and there are numerous random, uncaused events. (Morris, 1997, 19)

excerpts came from http://www.braungardt.com/Physics/Vacuum%20Fluctuation.htm

Friday, May 22, 2009

Sidhartha-The Buddha

Sidhartha was born (c. 563 BC; Kapilavastu, Nepal) into the Gautama family of the Shakaya clan. The Shakayas were members of the priestly-warrior caste. In fact, Sidhartha's father was the head of the tribe so Sidhartha was a prince and seemed destined to rule. He lived a luxurious life and the received the best education his father's wealth could provide, but his father also sheltered him from life's hardships.

He married a woman named Yashodhara and they lived in his father's house. Sidhartha was still protected from the trials of life. Yashodhara bore a son, and Sidhartha believed that he was happy.

Then, during one of his few excursions from the protection of his father's palace, Sidhartha saw three things which opened the harsh realities of life to him. He saw an old man, suffering from the frailties of age. He saw a sick man, suffering from disease. He also saw a dead man, which shocked him greatly. He finally realized that the infirmities of old age, and the pain of sickness and death caused suffering that he had never experienced. This revelation caused him to begin a search for truth that drastically changed his life, and, eventually, the lives of millions.


buddha.gif (57317 bytes)
An ancient sculpture of the Buddha

At the age of twenty-nine he left his home, his wife, his son, and his father. He gave up his claim to the succession of his father's throne and left the palace. He studied Yogic meditation with two Brahman hermits and achieved high cognitive states in both trance and meditation, but his desire for absolute truth was not satisfied.

For the next six years, Sidhartha placed his body under severe asceticism, which included extreme fasting and suspension of breathing. These practices almost killed him, but they did not satisfy his search for truth.

He finally ended his acetic lifestyle and began to eat. Sidhartha decided to meditate until the absolute truth would lie clearly in front of him. He meditated under a Bodhi tree where he sat facing east.

At the age of thirty-five, on the night of the full moon, Sidhartha reached enlightenment and became an "enlightened one"--a Buddha (c. 528 BC) He had at last discovered the truth he had sought, and he immediately shared it with five ascetics who had practiced near him.

After a few weeks of rest, he decided to teach the way to enlightenment to others and went to Deer Garden where he held his first sermon, "The turning wheel of Dharma." Sidhartha felt a strong call to teach others even though he could never teach the content of enlightenment, only the way of enlightenment. Buddha called his teachings "the middle way", because it was in the middle between asceticism and indulgence.

For the next forty-five years he taught as the Buddha or "Shakyamuni" (sage of the shakaya"). He also established a community of monks called sanga.

The Buddha died after forty-five years of teaching at the age of eighty.

excerpt taken from http://www.thenagain.info/WebChron/India/Buddha.html

Thursday, May 21, 2009

Natural Selection

By: Charles Darwin

IN order to make it clear how, as I believe, natural selection acts, I must beg permission to give one or two imaginary illustrations. Let us take the case of a wolf, which preys on various animals, securing some by craft, some by strength, and some by fleetness; and let us suppose that the fleetest prey, a deer for instance, had from any change in the country increased in numbers, or that other prey had decreased in numbers, during that season of the year when the wolf was hardest pressed for food. Under such circumstances the swiftest and slimmest wolves would have the best chance of surviving and so be preserved or selected,—provided always that they retained strength to master their prey at this or some other period of the year, when they were compelled to prey on other animals. I can see no more reason to doubt that this would be the result, than that man should be able to improve the fleetness of his greyhounds by careful and methodical selection, or by that kind of unconscious selection which follows from each man trying to keep the best dogs without any thought of modifying the breed. I may add, that, according to Mr. Pierce, there are two varieties of the wolf inhabiting the Catskill Mountains, in the United States, one with a light greyhound-like form, which pursues deer, and the other more bulky, with shorter legs, which more frequently attacks the shepherd’s flocks.

It should be observed that, in the above illustration, I speak of the slimmest individual wolves, and not of any single strongly-marked variation having been preserved. In former editions of this work I sometimes spoke as if this latter alternative had frequently occurred. I saw the great importance of individual differences, and this led me fully to discuss the results of unconscious selection by man, which depends on the preservation of all the more or less valuable individuals, and on the destruction of the worst. I saw, also, that the preservation in a state of nature of any occasional deviation of structure, such as a monstrosity, would be a rare event; and that, if at first preserved, it would generally be lost by subsequent intercrossing with ordinary individuals. Nevertheless, until reading an able and valuable article in the ‘North British Review’ (1867), I did not appreciate how rarely single variations, whether slight or strongly-marked, could be. perpetuated. The author takes the case of a pair of animals, producing during their lifetime two hundred offspring, of which, from various causes of destruction, only two on an average survive to procreate their kind. This is rather an extreme estimate for most of the higher animals, but by no means so for many of the lower organisms. He then shows that if a single individual were born, which varied in some manner, giving it twice as good a chance of life as that of the other individuals, yet the chances would be strongly against its survival. Supposing it to survive and to breed, and that half its young inherited the favourable variation; still, as the reviewer goes on to show, the young would have only a slightly better chance of surviving and breeding; and this chance would go on decreasing in the succeeding generations. The justice of these remarks cannot, I think, be disputed. If, for instance, a bird of some kind could procure its food more easily by having its beak curved, and if one were born with its beak strongly curved, and which consequently flourished, nevertheless there would be a very poor chance of this one individual perpetuating its kind to the exclusion of the common form; but there can hardly be a doubt, judging by what we see taking place under domestication, that this result would follow from the preservation during many generations of a large number of individuals with more or less strongly curved beaks, and from the destruction of a still larger number with the straightest beaks.

It should not, however, be overlooked that certain rather strongly marked variations, which no one would rank as mere individual differences, frequently recur owing to a similar organisation being similarly acted on—of which fact numerous instances could be given with our domestic productions. In such cases, if the varying individual did not actually transmit to its offspring its newly-acquired character, it would undoubtedly transmit to them, as long as the existing conditions remained the same, a still stronger tendency to vary in the same manner. There can also be little doubt that the tendency to vary in the same manner has often been so strong that all the individuals of the same species have been similarly modified without the aid of any form of selection. Or only a third, fifth, or tenth part of the individuals may have been thus affected, of which fact several instances could be given. Thus Graba estimates that about one-fifth of the guillemots in the Faroe Islands consist of a variety so well marked, that it was formerly ranked as a distinct species under the name of Uria lacrymans. In cases of this kind, if the variation were of a beneficial nature, the original form would soon be supplanted by the modified form, through the survival of the fittest.

To the effects of intercrossing in eliminating variations of all kinds, I shall have to recur; but it may be here remarked that most animals and plants keep to their proper homes, and do not needlessly wander about; we see this even with migratory birds, which almost always return to the same spot. Consequently each newly-formed variety would generally be at first local, as seems to be the common rule with varieties in a state of nature; so that similarly modified individuals would soon exist in a small body together, and would often breed together. If the new variety were successful in its battle for life, it would slowly spread from a central district, competing with and conquering the unchanged individuals on the margins of an ever-increasing circle.

It may be worth while to give another and more complex illustration of the action of natural selection. Certain plants excrete sweet juice, apparently for the sake of eliminating something injurious from the sap: this is effected, for instance, by glands at the base of the stipules in some Leguminosæ and at the backs of the leaves of the common laurel. This juice, though small in quantity, is greedily sought by insects; but their visits do not in any way benefit the plant. Now, let us suppose that the juice or nectar was excreted from the inside of the flowers of a certain number of plants of any species. Insects in seeking the nectar would get dusted with pollen, and would often transport it from one flower to another. The flowers of two distinct individuals of the same species would thus get crossed; and the act of crossing, as can be fully proved, gives rise to vigorous seedlings which consequently would have the best chance of flourishing and surviving The plants which produced flowers with the largest glands or nectaries, excreting most nectar, would oftenest be visited by insects, and would oftenest be crossed; and so in the long run would gain the upper hand and form a local variety. The flowers, also, which had their stamens and pistils placed, in relation to the size and habits of the particular insects which visited them, so as to favour in any degree the transportal of the pollen, would likewise be favoured. We might have taken the case of insects visiting flowers for the sake of collecting pollen instead of nectar; and as pollen is formed for the sole purpose of fertilisation, its destruction appears to be a simple loss to the plant; yet if a little pollen were carried, at first occasionally and then habitually, by the pollen-devouring insects from flower to flower, and a cross thus effected, although nine-tenths of the pollen were destroyed it might still be a great gain to the plant to be thus robbed; and the individuals which produced more and more pollen, and had larger anthers, would be selected.
When our plant, by the above process long continued, had been rendered highly attractive to insects, they would, unintentionally on their part, regularly carry pollen from flower to flower; and that they do this effectually, I could easily show by many striking facts. I will give only one, as likewise illustrating one step in the separation of the sexes of plants. Some holly trees bear only male flowers, which have four stamens producing a rather small quantity of pollen, and a rudimentary pistil; other holly trees bear only female flowers; these have a full-sized pistil, and four stamens with shrivelled anthers, in which not a grain of pollen can be detected. Having found a female tree exactly sixty yards from a male tree, I put the stigmas of twenty flowers, taken from different branches, under the microscope, and on all, without exception, there were a few pollen grains, and on some a profusion. As the wind had set for several days from the female to the male tree, the pollen could not thus have been carried. The weather had been cold and boisterous, and therefore not favourable to bees, nevertheless every female flower which I examined had been effectually fertilised by the bees, which had flown from tree to tree in search of nectar. But to return to our imaginary case: as soon as the plant had been rendered so highly attractive to insects that pollen was regularly carried from flower to flower, another process might commence. No naturalist doubts the advantage of what has been called the “physiological division of labour”; hence we may believe that it would be advantageous to a plant to produce stamens alone in one flower or on one whole plant, and pistils alone in another flower or on another plant. In plants under culture and placed under new conditions of life, sometimes the male organs and sometimes the female organs become more or less impotent; now if we suppose this to occur in ever so slight a degree under nature, then, as pollen is already carried regularly from flower to flower, and as a more complete separation of the sexes of our plant would be advantageous on the principle of the division of labour, individuals with this tendency more and more increased, would be continually favoured or selected, until at last a complete separation of the sexes might be effected. It would take up too much space to show the various steps, through dimorphism and other means, by which the separation of the sexes in plants of various kinds is apparently now in progress; but I may add that some of the species of holly in North America, are, according to Asa Gray, in an exactly intermediate condition, or, as he expresses it, are more or less dioeciously polygamous.
Let us now turn to the nectar-feeding insects; we may suppose the plant, of which we have been slowly increasing the nectar by continued selection, to be a common plant; and that certain insects depended in main part on its nectar for food. I could give many facts showing how anxious bees are to save time: for instance, their habit of cutting holes and sucking the nectar at the bases of certain flowers, which, with a very little more trouble, they can enter by the mouth. Bearing such facts in mind, it may be believed that under certain circumstances individual differences in the curvature or length of the proboscis, &c., too slight to be appreciated by us, might profit a bee or other insect, so that certain individuals would be able to obtain their food more quickly than others; and thus the communities to which they belonged would flourish and throw off many swarms inheriting the same peculiarities. The tubes of the corolla of the common red and incarnate clovers (Trifolium pratense and incarnatum) do not on a hasty glance appear to differ in length; yet the hive bee can easily suck the nectar out of the incarnate clover, but not out of the common red clover, which is visited by humble bees alone; so that whole fields of red clover offer in vain an abundant supply of precious nectar to the hive-bee. That this nectar is much liked by the hive bee is certain; for I have repeatedly seen, but only in the autumn, many hive bees sucking the flowers through holes bitten in the base of the tube by humble bees. The difference in the length of the corolla in the two kinds of clover, which determines the visits of the hive bee, must be very trifling; for I have been assured that when red clover has been mown, the flowers of the second crop are somewhat smaller, and that these are visited by many hive bees. I do not know whether this statement is accurate; nor whether another published statement can be trusted, namely, that the Ligurian bee which is generally considered a mere variety of the common hive bee, and which freely crosses with it, is able to reach and suck the nectar of the red clover. Thus, in a country where this kind of clover abounded, it might be a great advantage to the hive bee to have a slightly longer or differently constructed proboscis. On the other hand, as the fertility of this clover absolutely depends on bees visiting the flowers, if humble-bees were to become rare in any country, it might be a great advantage to the plant to have a, shorter or more deeply divided corolla, so that the hive bees should be enabled to suck its flowers. Thus I can understand how a flower and a bee might slowly become, either simultaneously or one after the other, modified and adapted to each other in the most perfect manner, by the continued preservation of all the individuals which presented slight deviations of structure mutually favourable to each other.
I am well aware that this doctrine of natural selection, exemplified in the above imaginary instances, is open to the same objections which were first urged against Sir Charles Lyell’s noble views on “the modern changes of the earth, as illustrative of geology”; but we now seldom hear the agencies which we see still at work, spoken of as trifling or insignificant, when used in explaining the excavation of the deepest valleys or the formation of long lines of inland cliffs. Natural selection acts only by the preservation and accumulation of small inherited modifications, each profitable to the preserved being; and as modern geology has almost banished such views as the excavation of a great valley by a single diluvial wave, so will natural selection banish the belief of the continued creation of new organic beings, or of any great and sudden modification in their structure.

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